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Myo15a myosin XVA [ Mus musculus (house mouse) ]

Gene ID: 17910, updated on 2-Nov-2024

Summary

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Official Symbol
Myo15aprovided by MGI
Official Full Name
myosin XVAprovided by MGI
Primary source
MGI:MGI:1261811
See related
Ensembl:ENSMUSG00000042678 AllianceGenome:MGI:1261811
Gene type
protein coding
RefSeq status
VALIDATED
Organism
Mus musculus
Lineage
Eukaryota; Metazoa; Chordata; Craniata; Vertebrata; Euteleostomi; Mammalia; Eutheria; Euarchontoglires; Glires; Rodentia; Myomorpha; Muroidea; Muridae; Murinae; Mus; Mus
Also known as
sh2; sh-2; Myo15
Summary
Predicted to enable actin filament binding activity and microfilament motor activity. Acts upstream of or within inner ear morphogenesis; locomotory behavior; and sensory perception of sound. Located in stereocilium. Is expressed in several structures, including brain; heart; inner ear; liver; and metanephros. Used to study autosomal recessive nonsyndromic deafness 3. Human ortholog(s) of this gene implicated in autosomal recessive nonsyndromic deafness 3 and sensorineural hearing loss. Orthologous to human MYO15A (myosin XVA). [provided by Alliance of Genome Resources, Nov 2024]
Expression
Biased expression in testis adult (RPKM 2.4), cerebellum adult (RPKM 0.5) and 4 other tissues See more
Orthologs
human all
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Genomic context

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See Myo15a in Genome Data Viewer
Location:
11 B2; 11 37.81 cM
Exon count:
67
Annotation release Status Assembly Chr Location
RS_2024_02 current GRCm39 (GCF_000001635.27) 11 NC_000077.7 (60360165..60419195)
108.20200622 previous assembly GRCm38.p6 (GCF_000001635.26) 11 NC_000077.6 (60469339..60528369)

Chromosome 11 - NC_000077.7Genomic Context describing neighboring genes Neighboring gene CapStarr-seq enhancer MGSCv37_chr11:60267901-60268133 Neighboring gene ribosomal protein S4, X-linked pseudogene Neighboring gene developmentally regulated GTP binding protein 2 Neighboring gene predicted gene, 51901 Neighboring gene CapStarr-seq enhancer MGSCv37_chr11:60350718-60350827 Neighboring gene predicted gene, 53694 Neighboring gene alkB homolog 5, RNA demethylase Neighboring gene ubiquitin-60S ribosomal protein L40 pseudogene

Genomic regions, transcripts, and products

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Go to nucleotide: Graphics FASTA GenBank

Expression

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  • Project title: Mouse ENCODE transcriptome data Mouse ENCODE transcriptome data
  • Description: RNA profiling data sets generated by the Mouse ENCODE project.
  • BioProject: PRJNA66167
  • Publication: PMID 25409824
  • Analysis date: n/a

Bibliography

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Related articles in PubMed

  1. Incomplete maturation of brainstem auditory nuclei in genetically induced early postnatal cochlear degeneration. Webster DB, et al. Acta Otolaryngol, 1986 May-Jun. PMID 3727977
  2. Fast adaptation and Ca2+ sensitivity of the mechanotransducer require myosin-XVa in inner but not outer cochlear hair cells. Stepanyan R, et al. J Neurosci, 2009 Apr 1. PMID 19339598, Free PMC Article
  3. Auditory mechanotransduction in the absence of functional myosin-XVa. Stepanyan R, et al. J Physiol, 2006 Nov 1. PMID 16973713, Free PMC Article
  4. Transgene correction maintains normal cochlear structure and function in 6-month-old Myo15a mutant mice. Kanzaki S, et al. Hear Res, 2006 Apr. PMID 16580798
  5. Linkage Relations of Zavadskaia Shaker in the House Mouse (Mus musculus). Clark FH. Proc Natl Acad Sci U S A, 1935 May. PMID 16577664, Free PMC Article

See all (91) citations in PubMed

GeneRIFs: Gene References Into Functions

What's a GeneRIF?
  1. The ATPase mechanism of myosin 15, the molecular motor mutated in DFNB3 human deafness.
    Title: The ATPase mechanism of myosin 15, the molecular motor mutated in DFNB3 human deafness.
  2. Myosin-XVa Controls Both Staircase Architecture and Diameter Gradation of Stereocilia Rows in the Auditory Hair Cell Bundles.
    Title: Myosin-XVa Controls Both Staircase Architecture and Diameter Gradation of Stereocilia Rows in the Auditory Hair Cell Bundles.
  3. Gpsm2 defines an approximately 200 nm nanodomain at the tips of stereocilia and this localization requires the presence of Galphai3, myosin 15 and whirlin.
    Title: Defective Gpsm2/Gα(i3) signalling disrupts stereocilia development and growth cone actin dynamics in Chudley-McCullough syndrome.
  4. Myosin 15 isoforms can navigate between functionally distinct classes of stereocilia, and are independently required to assemble and then maintain the intricate hair bundle architecture.
    Title: The 133-kDa N-terminal domain enables myosin 15 to maintain mechanotransducing stereocilia and is essential for hearing.
  5. myo15-subfragment 1 is a barbed-end-directed motor that moves actin filaments in a gliding assay using a power stroke of 7.9 nm. Myo15-S1 was rate limited by transit through strongly actin-bound states, similar to other processive myosin motors.
    Title: Chaperone-enhanced purification of unconventional myosin 15, a molecular motor specialized for stereocilia protein trafficking.
  6. Genetic association between myoXV and deafness in a murine model.
    Title: Spontaneous mutant ICR kuru2 might be another shaker-2 deaf mouse.
  7. It was concluded that MyoXVa, whirlin, and Eps8 are integral components of the stereocilia tip complex, where Eps8 is a central actin-regulatory element for elongation of the stereocilia actin core.
    Title: Regulation of stereocilia length by myosin XVa and whirlin depends on the actin-regulatory protein Eps8.
  8. The Ca(2+) sensitivity of the mechanotransduction channels and the fast adaptation require a structural environment that is dependent on myosin-XVa and is disrupted in Myo15(sh2/sh2) inner hair cells, but not in Myo15(sh2/sh2) outer hair cells.
    Title: Fast adaptation and Ca2+ sensitivity of the mechanotransducer require myosin-XVa in inner but not outer cochlear hair cells.
  9. The morphology of cochlear hair cell stereocilia in double mutants reflects a dominance of the more severe Myo15 (sh2/sh2) phenotype over the Whrn (wi/wi) phenotype.
    Title: Whirler mutant hair cells have less severe pathology than shaker 2 or double mutants.
  10. This comparison revealed that the absence of functional myosin-XVa does not disrupt adaptation of the mechanotransduction current during sustained bundle deflection.
    Title: Auditory mechanotransduction in the absence of functional myosin-XVa.
Submit: New GeneRIF Correction See all GeneRIFs (12)

Variation

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Alleles

Alleles of this type are documented at Mouse Genome Informatics  (MGI)
  • Chemically induced (ENU) (4)  1 citation
  • Endonuclease-mediated (3) 
  • Radiation induced (1) 
  • Spontaneous (2) 
  • Targeted (3)  1 citation

Interactions

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Products Interactant Other Gene Complex Source Pubs Description

General gene information

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Markers

Homology

Gene Ontology Provided by MGI

Function Evidence Code Pubs
enables ATP binding IEA
Inferred from Electronic Annotation
more info
  
enables actin binding IEA
Inferred from Electronic Annotation
more info
  
enables actin filament binding IBA
Inferred from Biological aspect of Ancestor
more info
  
enables cytoskeletal motor activity IEA
Inferred from Electronic Annotation
more info
  
enables microfilament motor activity IBA
Inferred from Biological aspect of Ancestor
more info
  
enables protein binding IPI
Inferred from Physical Interaction
more info
 PubMed 
Process Evidence Code Pubs
involved_in actin filament organization IBA
Inferred from Biological aspect of Ancestor
more info
  
involved_in endocytosis IBA
Inferred from Biological aspect of Ancestor
more info
  
acts_upstream_of_or_within inner ear morphogenesis IMP
Inferred from Mutant Phenotype
more info
 PubMed 
acts_upstream_of_or_within locomotory behavior IMP
Inferred from Mutant Phenotype
more info
 PubMed 
involved_in response to light stimulus IEA
Inferred from Electronic Annotation
more info
  
involved_in response to light stimulus ISO
Inferred from Sequence Orthology
more info
  
acts_upstream_of_or_within sensory perception of sound IMP
Inferred from Mutant Phenotype
more info
 PubMed 
Component Evidence Code Pubs
is_active_in actin cytoskeleton IBA
Inferred from Biological aspect of Ancestor
more info
  
is_active_in actin-based cell projection IBA
Inferred from Biological aspect of Ancestor
more info
  
is_active_in cytoplasm IBA
Inferred from Biological aspect of Ancestor
more info
  
located_in cytosol TAS
Traceable Author Statement
more info
  
is_active_in membrane IBA
Inferred from Biological aspect of Ancestor
more info
  
part_of myosin complex IEA
Inferred from Electronic Annotation
more info
  
located_in stereocilium IDA
Inferred from Direct Assay
more info
 PubMed 
located_in stereocilium bundle IDA
Inferred from Direct Assay
more info
 PubMed 

General protein information

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Preferred Names
unconventional myosin-XV
Names
shaker 2
unconventional myosin-15

NCBI Reference Sequences (RefSeq)

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RefSeqs maintained independently of Annotated Genomes

These reference sequences exist independently of genome builds. Explain

These reference sequences are curated independently of the genome annotation cycle, so their versions may not match the RefSeq versions in the current genome build. Identify version mismatches by comparing the version of the RefSeq in this section to the one reported in Genomic regions, transcripts, and products above.

mRNA and Protein(s)

  1. NM_001103171.1 → NP_001096641.1  unconventional myosin-XV isoform 3

    Status: VALIDATED

    Description
    Transcript Variant: This variant (3) lacks an in-frame exon in the 5' coding region, compared to variant 1, resulting in a shorter protein (isoform 3), compared to isoform 1.
    Source sequence(s)
    AL596090, AL596386, AY331132
    Consensus CDS
    CCDS48811.1
    UniProtKB/Swiss-Prot
    Q9QZZ4
    Related
    ENSMUSP00000091686.3, ENSMUST00000094135.9
    Conserved Domains (7) summary
    smart00139
    Location:3013 → 3167
    MyTH4; Domain in Myosin and Kinesin Tails
    smart00242
    Location:1201 → 1883
    MYSc; Myosin. Large ATPases
    smart00295
    Location:3177 → 3382
    B41; Band 4.1 homologues
    cd13201
    Location:3378 → 3479
    FERM_C_MyoXV; FERM domain C-lobe of Myosin XV (MyoXV/Myo15)
    cd14473
    Location:3272 → 3372
    FERM_B-lobe; FERM domain B-lobe
    cd01387
    Location:1220 → 1871
    MYSc_Myo15; class XV mammal-like myosin, motor domain
    cl17036
    Location:2834 → 2913
    SH3; Src Homology 3 domain superfamily

  2. NM_010862.2 → NP_034992.2  unconventional myosin-XV isoform 1

    See identical proteins and their annotated locations for NP_034992.2

    Status: VALIDATED

    Description
    Transcript Variant: This variant (1) represents the longest transcript and encodes the longest isoform (1).
    Source sequence(s)
    AF144095, AL596090, AL596386
    Consensus CDS
    CCDS24792.1
    UniProtKB/Swiss-Prot
    A2A637, A2A638, O70395, Q5SX93, Q7TMR5, Q7TMR6, Q9QWL6, Q9QZZ4
    Related
    ENSMUSP00000071777.3, ENSMUST00000071880.9
    Conserved Domains (8) summary
    smart00139
    Location:3031 → 3185
    MyTH4; Domain in Myosin and Kinesin Tails
    smart00242
    Location:1201 → 1883
    MYSc; Myosin. Large ATPases
    smart00295
    Location:3195 → 3400
    B41; Band 4.1 homologues
    cd13201
    Location:3396 → 3497
    FERM_C_MyoXV; FERM domain C-lobe of Myosin XV (MyoXV/Myo15)
    cd14473
    Location:3290 → 3390
    FERM_B-lobe; FERM domain B-lobe
    cd01387
    Location:1220 → 1871
    MYSc_Myo15; class XV mammal-like myosin, motor domain
    pfam02050
    Location:1870 → 1960
    FliJ; Flagellar FliJ protein
    cl17036
    Location:2852 → 2931
    SH3; Src Homology 3 domain superfamily

  3. NM_182698.2 → NP_874357.2  unconventional myosin-XV isoform 2a

    Status: VALIDATED

    Description
    Transcript Variant: This variant (2) lacks two in-frame exons in the 5' coding region, compared to variant 1, resulting in a protein (isoform 2a) with a shorter N-terminus, compared to isoform 1.
    Source sequence(s)
    AL596386, AY331133
    Consensus CDS
    CCDS24793.1
    UniProtKB/Swiss-Prot
    Q9QZZ4
    Related
    ENSMUSP00000080507.6, ENSMUST00000081823.12
    Conserved Domains (7) summary
    smart00139
    Location:1826 → 1980
    MyTH4; Domain in Myosin and Kinesin Tails
    smart00242
    Location:14 → 696
    MYSc; Myosin. Large ATPases
    smart00295
    Location:1990 → 2195
    B41; Band 4.1 homologues
    cd13201
    Location:2191 → 2292
    FERM_C_MyoXV; FERM domain C-lobe of Myosin XV (MyoXV/Myo15)
    cd14473
    Location:2085 → 2185
    FERM_B-lobe; FERM domain B-lobe
    cd01387
    Location:33 → 684
    MYSc_Myo15; class XV mammal-like myosin, motor domain
    cl17036
    Location:1647 → 1726
    SH3; Src Homology 3 domain superfamily

RefSeqs of Annotated Genomes: GCF_000001635.27-RS_2024_02

The following sections contain reference sequences that belong to a specific genome build. Explain

Reference GRCm39 C57BL/6J

Genomic

  1. NC_000077.7 Reference GRCm39 C57BL/6J

    Range
    60360165..60419195
    Download
    GenBank, FASTA, Sequence Viewer (Graphics)

mRNA and Protein(s)

  1. XM_017314331.2 → XP_017169820.1  unconventional myosin-XV isoform X13

    UniProtKB/TrEMBL
    F6VXK7
    Related
    ENSMUSP00000120839.2, ENSMUST00000126522.4
    Conserved Domains (5) summary
    smart00015
    Location:632 → 653
    IQ; Calmodulin-binding motif
    smart00139
    Location:772 → 918
    MyTH4; Domain in Myosin and Kinesin Tails
    PHA03247
    Location:1139 → 1377
    PHA03247; large tegument protein UL36; Provisional
    cl22853
    Location:53 → 594
    Motor_domain; Myosin and Kinesin motor domain
    cl17036
    Location:1587 → 1666
    SH3; Src Homology 3 domain superfamily

  2. XM_017314330.2 → XP_017169819.1  unconventional myosin-XV isoform X12

    UniProtKB/TrEMBL
    F6VXK7
    Conserved Domains (5) summary
    smart00015
    Location:632 → 653
    IQ; Calmodulin-binding motif
    smart00139
    Location:772 → 918
    MyTH4; Domain in Myosin and Kinesin Tails
    PHA03247
    Location:1139 → 1377
    PHA03247; large tegument protein UL36; Provisional
    cl22853
    Location:53 → 594
    Motor_domain; Myosin and Kinesin motor domain
    cl17036
    Location:1587 → 1666
    SH3; Src Homology 3 domain superfamily

  3. XM_017314329.2 → XP_017169818.1  unconventional myosin-XV isoform X11

    UniProtKB/TrEMBL
    F6VXK7
    Conserved Domains (5) summary
    smart00015
    Location:632 → 653
    IQ; Calmodulin-binding motif
    smart00139
    Location:1766 → 1920
    MyTH4; Domain in Myosin and Kinesin Tails
    PHA03247
    Location:1139 → 1377
    PHA03247; large tegument protein UL36; Provisional
    cl22853
    Location:53 → 594
    Motor_domain; Myosin and Kinesin motor domain
    cl17036
    Location:1587 → 1666
    SH3; Src Homology 3 domain superfamily

  4. XM_017314324.3 → XP_017169813.1  unconventional myosin-XV isoform X6

    Conserved Domains (7) summary
    smart00015
    Location:632 → 653
    IQ; Calmodulin-binding motif
    smart00139
    Location:1739 → 1893
    MyTH4; Domain in Myosin and Kinesin Tails
    PHA03247
    Location:1164 → 1350
    PHA03247; large tegument protein UL36; Provisional
    cd13201
    Location:2104 → 2205
    FERM_C_MyoXV; FERM domain C-lobe of Myosin XV (MyoXV/Myo15)
    cd14473
    Location:1998 → 2098
    FERM_B-lobe; FERM domain B-lobe
    cl22853
    Location:53 → 594
    Motor_domain; Myosin and Kinesin motor domain
    cl17036
    Location:1560 → 1639
    SH3; Src Homology 3 domain superfamily

  5. XM_017314320.3 → XP_017169809.1  unconventional myosin-XV isoform X2

    Conserved Domains (7) summary
    smart00015
    Location:632 → 653
    IQ; Calmodulin-binding motif
    smart00139
    Location:1766 → 1920
    MyTH4; Domain in Myosin and Kinesin Tails
    PHA03247
    Location:1139 → 1377
    PHA03247; large tegument protein UL36; Provisional
    cd13201
    Location:2131 → 2232
    FERM_C_MyoXV; FERM domain C-lobe of Myosin XV (MyoXV/Myo15)
    cd14473
    Location:2025 → 2125
    FERM_B-lobe; FERM domain B-lobe
    cl22853
    Location:53 → 594
    Motor_domain; Myosin and Kinesin motor domain
    cl17036
    Location:1587 → 1666
    SH3; Src Homology 3 domain superfamily

  6. XM_017314319.3 → XP_017169808.1  unconventional myosin-XV isoform X1

    Conserved Domains (7) summary
    smart00015
    Location:632 → 653
    IQ; Calmodulin-binding motif
    smart00139
    Location:1766 → 1920
    MyTH4; Domain in Myosin and Kinesin Tails
    PHA03247
    Location:1139 → 1377
    PHA03247; large tegument protein UL36; Provisional
    cd13201
    Location:2131 → 2232
    FERM_C_MyoXV; FERM domain C-lobe of Myosin XV (MyoXV/Myo15)
    cd14473
    Location:2025 → 2125
    FERM_B-lobe; FERM domain B-lobe
    cl22853
    Location:53 → 594
    Motor_domain; Myosin and Kinesin motor domain
    cl17036
    Location:1587 → 1666
    SH3; Src Homology 3 domain superfamily

  7. XM_017314328.3 → XP_017169817.1  unconventional myosin-XV isoform X10

    Conserved Domains (7) summary
    smart00015
    Location:632 → 653
    IQ; Calmodulin-binding motif
    smart00139
    Location:1704 → 1857
    MyTH4; Domain in Myosin and Kinesin Tails
    PHA03247
    Location:1139 → 1377
    PHA03247; large tegument protein UL36; Provisional
    cd13201
    Location:2068 → 2169
    FERM_C_MyoXV; FERM domain C-lobe of Myosin XV (MyoXV/Myo15)
    cd14473
    Location:1962 → 2062
    FERM_B-lobe; FERM domain B-lobe
    cl22853
    Location:53 → 594
    Motor_domain; Myosin and Kinesin motor domain
    cl17036
    Location:1587 → 1666
    SH3; Src Homology 3 domain superfamily

  8. XM_017314321.3 → XP_017169810.1  unconventional myosin-XV isoform X3

    Conserved Domains (7) summary
    smart00015
    Location:632 → 653
    IQ; Calmodulin-binding motif
    smart00139
    Location:1765 → 1919
    MyTH4; Domain in Myosin and Kinesin Tails
    PHA03247
    Location:1138 → 1376
    PHA03247; large tegument protein UL36; Provisional
    cd13201
    Location:2130 → 2231
    FERM_C_MyoXV; FERM domain C-lobe of Myosin XV (MyoXV/Myo15)
    cd14473
    Location:2024 → 2124
    FERM_B-lobe; FERM domain B-lobe
    cl22853
    Location:53 → 594
    Motor_domain; Myosin and Kinesin motor domain
    cl17036
    Location:1586 → 1665
    SH3; Src Homology 3 domain superfamily

  9. XM_017314327.3 → XP_017169816.1  unconventional myosin-XV isoform X9

    Conserved Domains (8) summary
    smart00015
    Location:632 → 653
    IQ; Calmodulin-binding motif
    smart00139
    Location:1723 → 1877
    MyTH4; Domain in Myosin and Kinesin Tails
    PHA03247
    Location:1096 → 1334
    PHA03247; large tegument protein UL36; Provisional
    cd13201
    Location:2088 → 2189
    FERM_C_MyoXV; FERM domain C-lobe of Myosin XV (MyoXV/Myo15)
    cd14473
    Location:1982 → 2082
    FERM_B-lobe; FERM domain B-lobe
    cl22853
    Location:53 → 594
    Motor_domain; Myosin and Kinesin motor domain
    cl02480
    Location:772 → 875
    MyTH4; MyTH4 domain
    cl17036
    Location:1544 → 1623
    SH3; Src Homology 3 domain superfamily

  10. XM_017314323.3 → XP_017169812.1  unconventional myosin-XV isoform X5

    Conserved Domains (7) summary
    smart00015
    Location:632 → 653
    IQ; Calmodulin-binding motif
    smart00139
    Location:1748 → 1902
    MyTH4; Domain in Myosin and Kinesin Tails
    PHA03247
    Location:1121 → 1359
    PHA03247; large tegument protein UL36; Provisional
    cd13201
    Location:2113 → 2214
    FERM_C_MyoXV; FERM domain C-lobe of Myosin XV (MyoXV/Myo15)
    cd14473
    Location:2007 → 2107
    FERM_B-lobe; FERM domain B-lobe
    cl22853
    Location:53 → 594
    Motor_domain; Myosin and Kinesin motor domain
    cl17036
    Location:1569 → 1648
    SH3; Src Homology 3 domain superfamily

  11. XM_017314326.3 → XP_017169815.1  unconventional myosin-XV isoform X8

    Conserved Domains (7) summary
    smart00015
    Location:594 → 615
    IQ; Calmodulin-binding motif
    smart00139
    Location:1728 → 1882
    MyTH4; Domain in Myosin and Kinesin Tails
    PHA03247
    Location:1101 → 1339
    PHA03247; large tegument protein UL36; Provisional
    cd13201
    Location:2093 → 2194
    FERM_C_MyoXV; FERM domain C-lobe of Myosin XV (MyoXV/Myo15)
    cd14473
    Location:1987 → 2087
    FERM_B-lobe; FERM domain B-lobe
    cl22853
    Location:53 → 556
    Motor_domain; Myosin and Kinesin motor domain
    cl17036
    Location:1549 → 1628
    SH3; Src Homology 3 domain superfamily

  12. XM_017314325.3 → XP_017169814.1  unconventional myosin-XV isoform X7

    Conserved Domains (7) summary
    smart00015
    Location:594 → 615
    IQ; Calmodulin-binding motif
    smart00139
    Location:1728 → 1882
    MyTH4; Domain in Myosin and Kinesin Tails
    PHA03247
    Location:1101 → 1339
    PHA03247; large tegument protein UL36; Provisional
    cd13201
    Location:2093 → 2194
    FERM_C_MyoXV; FERM domain C-lobe of Myosin XV (MyoXV/Myo15)
    cd14473
    Location:1987 → 2087
    FERM_B-lobe; FERM domain B-lobe
    cl22853
    Location:53 → 556
    Motor_domain; Myosin and Kinesin motor domain
    cl17036
    Location:1549 → 1628
    SH3; Src Homology 3 domain superfamily

  13. XM_017314322.3 → XP_017169811.1  unconventional myosin-XV isoform X4

    Conserved Domains (7) summary
    smart00015
    Location:632 → 653
    IQ; Calmodulin-binding motif
    smart00139
    Location:1755 → 1909
    MyTH4; Domain in Myosin and Kinesin Tails
    PHA03247
    Location:1139 → 1377
    PHA03247; large tegument protein UL36; Provisional
    cd13201
    Location:2120 → 2221
    FERM_C_MyoXV; FERM domain C-lobe of Myosin XV (MyoXV/Myo15)
    cd14473
    Location:2014 → 2114
    FERM_B-lobe; FERM domain B-lobe
    cl22853
    Location:53 → 594
    Motor_domain; Myosin and Kinesin motor domain
    cl17036
    Location:1576 → 1655
    SH3; Src Homology 3 domain superfamily

RNA

  1. XR_001779911.2 RNA Sequence

Nucleotide Protein Strain
Heading Accession and Version
Protein Accession Links
GenPept Link UniProtKB Link
Q9QZZ4.2 GenPept UniProtKB/Swiss-Prot:Q9QZZ4

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